By default a new message has only two fields - 'To' and 'Subject' and it's perfectly enough for 90% cases when you have your reply-to address and signature pre-defined in your email account settings and basically you don't need to change anything in your message header fields.
Now when want to change signature for different messages, reply-to address or even smtp server to send a message from, you need a bit of customization and here is what you need to do:
- Click on the small square field in front of a Subject line and click on 'Customize' and now by checking these fields you can specify which fields you want to have in hand for all your new messages.
- CC, BCC, Reply-to and Priority fields are on the list next to 'Customize' item on the same pop-up menu.
To specify different signatures you need go to 'Preferences --> Signatures', to specify different smtp servers to send your messages from go to 'Preferences --> Accounts --> Choose account you want to specify servers for and then click on drop down list at 'Outgoing Mail Server (SMTP): ---> Edit Server List.


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